:root{--ink:#073e42;--lime:#a5be32;--paper:#f7f5ef;--muted:#526767;--line:#d8ddd3}*{box-sizing:border-box}html{scroll-behavior:smooth;scroll-padding-top:100px}body{margin:0;background:var(--paper);color:var(--ink);font-family:"Helvetica Neue",Helvetica,Arial,sans-serif;font-size:17px;line-height:1.6}a{color:inherit;text-decoration:none}a:hover{text-decoration:underline;text-underline-offset:5px}a:focus-visible,summary:focus-visible{outline:3px solid #7c9420;outline-offset:6px}button,a{-webkit-tap-highlight-color:transparent}.wrap{max-width:1440px;margin:auto;padding-left:6%;padding-right:6%}.header{display:flex;justify-content:space-between;align-items:center;gap:24px;padding:18px 6%;border-bottom:1px solid var(--line);max-width:1600px;margin:auto}.logo{width:245px;height:auto;display:block}.header nav{display:flex;align-items:center;gap:30px;font-size:15px}.nav-contact{padding:10px 18px;border:1px solid var(--ink);border-radius:4px}.nav-contact span{margin-left:16px}.hero{padding-top:76px;display:grid;grid-template-columns:1.25fr 1fr;gap:30px;align-items:center}.eyebrow{text-transform:uppercase;letter-spacing:.15em;font-size:13px;font-weight:600;margin:0 0 26px}.hero h1{font-size:clamp(44px,5.2vw,78px);line-height:1.06;font-weight:500;letter-spacing:-.055em;margin:0 0 30px}.hero h1 em{font-style:normal;color:#73891f}.intro{max-width:560px;font-size:23px;line-height:1.45;letter-spacing:-.02em}.hero-detail{max-width:465px;color:var(--muted);margin:20px 0 30px}.button{display:inline-flex;align-items:center;gap:32px;background:var(--ink);color:var(--paper);padding:16px 23px;border-radius:4px;font-size:16px;font-weight:500;transition:background .2s,transform .2s}.button:hover{background:#14585b;text-decoration:none;transform:translateY(-2px)}.hero-art{position:relative;padding:32px 18px 20px;border-left:1px solid var(--line)}.hero-art img{width:100%;height:auto;display:block;max-height:420px}.art-label{display:block;text-align:center;font-size:14px;color:var(--muted)}.art-caption{display:flex;justify-content:space-between;gap:20px;font-size:12px;letter-spacing:.02em;border-top:1px solid var(--line);padding-top:18px}.hero-footer{grid-column:1/-1;border-top:1px solid var(--line);padding:23px 0;margin-top:35px;display:flex;justify-content:space-between;font-size:14px}.muted{color:var(--muted)}.footer{display:flex;gap:28px;justify-content:space-between;padding-top:30px;padding-bottom:30px;border-top:1px solid var(--line);font-size:13px}.skip{position:absolute;left:10px;top:-100px;background:var(--ink);color:white;padding:12px;z-index:10}.skip:focus{top:10px}section{scroll-margin-top:30px}@media(max-width:900px){.header nav{gap:18px}.hero{padding-top:50px;grid-template-columns:1.2fr 1fr}.hero-art{padding:20px 0 20px 15px}.art-caption{display:block}.art-caption span{display:block}.intro{font-size:20px}.logo{width:215px}}@media(max-width:680px){.header{padding:12px 5%;flex-wrap:wrap;gap:6px}.logo{width:220px}.header nav{width:100%;justify-content:space-between;gap:12px;font-size:14px}.nav-contact{padding:6px 10px}.nav-contact span{margin-left:6px}.wrap{padding-left:6%;padding-right:6%}.hero{grid-template-columns:1fr;gap:20px;padding-top:45px}.hero h1{font-size:clamp(42px,10vw,64px)}.hero-art{border-left:0;padding:10px 30px;max-width:400px;width:100%;margin:auto}.hero-art img{max-height:300px}.art-caption{display:flex;font-size:12px}.hero-footer{margin-top:10px;display:block;font-size:13px}.hero-footer>span{display:block}.footer{flex-wrap:wrap;gap:12px}.footer>span:last-child{width:100%}}@media(prefers-reduced-motion:reduce){html{scroll-behavior:auto}.button{transition:none}}
.section{padding-top:100px;padding-bottom:100px}.section-heading{max-width:760px;margin-bottom:54px}h2{font-size:clamp(36px,3.7vw,56px);font-weight:500;letter-spacing:-.045em;line-height:1.12;margin:0 0 25px}.section-heading>p:last-child{max-width:560px;color:var(--muted);font-size:19px}h3{font-size:25px;letter-spacing:-.025em;font-weight:500;line-height:1.25;margin:0 0 18px}.services{display:grid;grid-template-columns:repeat(2,minmax(0,1fr));gap:54px 60px;align-items:start}.service{border-top:2px solid var(--ink);padding-top:24px}.service-number{display:block;font-size:14px;color:#74881f;margin-bottom:36px}.service p{color:var(--muted);font-size:16px}.service .question{color:var(--ink);font-size:18px;min-height:58px;line-height:1.5}.tags{padding:0;list-style:none;display:flex;flex-wrap:wrap;gap:8px;margin:26px 0}.tags li{border:1px solid #d4d9cb;padding:5px 10px;border-radius:3px;font-size:13px;line-height:1.5}.service .deliverable{border-top:1px solid var(--line);padding-top:20px;margin:0}.deliverable strong{display:block;color:var(--ink);font-size:13px;letter-spacing:.04em;margin-bottom:8px}.experience{background:var(--ink);color:var(--paper);padding:90px 0 35px}.experience .eyebrow{color:#c0d865}.experience .section-heading>p:last-child{color:#bed1cf}.work-grid{display:grid;grid-template-columns:repeat(3,1fr);gap:44px}.work-grid article{border-top:1px solid #43676a;padding-top:24px}.work-context{font-size:13px;color:#c0d865;margin:0 0 28px;min-height:42px}.work-grid h3{font-size:26px;line-height:1.3}.work-grid article>p:not(.work-context){color:#c6d5d3;font-size:16px;line-height:1.7}.work-result{border-left:2px solid var(--lime);padding-left:16px;font-size:15px;margin-top:28px}.experience-note{font-size:12px;color:#b2c7c5;margin-top:50px}.engagement{display:grid;grid-template-columns:1fr 1.15fr;gap:11%}.engagement>div>p:not(.eyebrow){color:var(--muted);max-width:390px}.text-link{display:inline-block;font-size:15px;font-weight:500;border-bottom:1px solid var(--ink);padding-bottom:5px;margin-top:20px}.text-link span{margin-left:20px}.text-link:hover{text-decoration:none;border-bottom-width:2px}.engagement-list article{display:flex;gap:28px;padding:28px 0;border-top:1px solid var(--line)}.engagement-list article:first-child{padding-top:0;border-top:0}.engagement-list article>span{font-size:13px;margin-top:5px;color:#73891f}.engagement-list h3{font-size:23px;margin-bottom:10px}.engagement-list p{font-size:16px;color:var(--muted);margin:0}.about{border-top:1px solid var(--line);padding-top:80px;padding-bottom:95px;display:grid;grid-template-columns:1fr 1.15fr;gap:11%}.about h2{font-size:46px;margin-bottom:12px}.credentials{font-size:19px;margin:0 0 40px}.about-location{font-size:14px;color:var(--muted)}.large-copy{font-size:28px;letter-spacing:-.025em;line-height:1.35;margin:0 0 24px}.about-copy>p:not(.large-copy){color:var(--muted);font-size:17px}.about-facts{display:grid;grid-template-columns:1fr 1fr;gap:20px;margin-top:34px;padding-top:23px;border-top:1px solid var(--line);font-size:14px}.about-facts strong,.about-facts span{display:block}.about-facts span{color:var(--muted);margin-top:5px}.contact{background:var(--lime);padding:70px 0}.contact-inner{display:flex;justify-content:space-between;align-items:center;gap:40px}.contact h2{font-size:clamp(34px,3.2vw,49px)}.contact p:not(.eyebrow){max-width:540px;font-size:18px}.contact-action{min-width:240px}.contact .email{display:block;margin-top:16px;font-size:16px}.footer{border:0}.footer>a{margin-left:auto}.logo{width:240px}.header{padding-top:25px;padding-bottom:25px}@media(max-width:1000px){.services,.work-grid{gap:25px}.work-grid h3{font-size:23px}.engagement,.about{gap:7%}.contact-inner{align-items:flex-start}.contact-action{padding-top:50px}}@media(max-width:760px){.section{padding-top:65px;padding-bottom:65px}.section-heading{margin-bottom:35px}.services,.work-grid{grid-template-columns:1fr;gap:40px}.service{display:grid;grid-template-columns:30px 1fr;gap:20px}.service-number{margin-top:6px}.service .question{min-height:0}.service p{max-width:580px}.tags{margin:20px 0}.experience{padding-top:65px}.work-grid{gap:30px}.work-context{min-height:0;margin-bottom:18px}.work-result{margin-top:20px}.engagement,.about{grid-template-columns:1fr;gap:40px}.engagement>div>p:not(.eyebrow){max-width:100%}.about{padding-top:60px;padding-bottom:65px}.about h2{font-size:40px}.credentials{margin-bottom:14px}.large-copy{font-size:25px}.contact-inner{display:block}.contact-action{padding-top:15px}.contact{padding:55px 0}.header{padding-top:18px;padding-bottom:15px}.logo{width:215px;margin-bottom:10px}.about-facts{gap:15px}.footer>a{margin-left:0}}@media(max-width:400px){.art-caption{font-size:12px;gap:10px}.header nav{gap:8px;font-size:13px}.about-facts{grid-template-columns:1fr}.eyebrow{font-size:12px}}

/* Four service areas with progressive detail. */
.service-number{margin-bottom:22px;font-size:12px;letter-spacing:.1em;font-weight:600;color:#607516}
.service h3{font-size:clamp(24px,2.5vw,32px);margin-bottom:16px}
.service .service-intro{font-size:19px;color:var(--ink);line-height:1.45;margin:0 0 12px}
.service details{margin:24px 0}
.service summary{cursor:pointer;font-size:15px;font-weight:500;padding:10px 0;min-height:44px;list-style:none;display:flex;align-items:center;justify-content:space-between;gap:20px}
.service summary::-webkit-details-marker{display:none}
.service summary::after{content:'+';font-size:23px;font-weight:400;line-height:1}
.service details[open] summary::after{content:'−'}
.service summary:hover{color:#607516}
.capabilities{margin:10px 0 24px;padding-left:20px;font-size:16px;color:var(--muted)}
.capabilities li{padding-left:4px;margin-bottom:15px}
.capabilities strong{color:var(--ink);font-weight:500}
.service .deliverable{font-size:15px}
@media(max-width:760px){.services{grid-template-columns:1fr;gap:38px}.service{display:block}.service-number{margin-bottom:18px}.service h3{font-size:27px}.service details{margin:18px 0}}

/* Selected research, kept compact and linked to the published papers. */
.publications{border-top:1px solid var(--line);padding-top:75px}
.publication-heading{display:flex;justify-content:space-between;align-items:flex-end;gap:40px;margin-bottom:38px}
.publication-heading h2{margin-bottom:0}
.publication-heading>.text-link{margin-bottom:5px;flex-shrink:0;font-size:14px}
.publication{display:grid;grid-template-columns:230px minmax(0,1fr);gap:42px;border-top:1px solid var(--line);padding:30px 0}
.publication:last-child{border-bottom:1px solid var(--line)}
.publication-meta{display:flex;flex-direction:column;align-items:flex-start;gap:7px;font-size:14px;color:var(--muted)}
.publication-meta>span:first-child{font-weight:500;color:var(--ink)}
.publication-meta .authorship{font-size:12px;border:1px solid var(--line);border-radius:3px;padding:2px 8px;margin-top:5px}
.publication h3{font-size:21px;line-height:1.4;letter-spacing:-.015em;margin:0 0 12px}
.publication h3 span{white-space:nowrap}
.publication p{font-size:16px;color:var(--muted);margin:0;max-width:760px}
.publications-jump{margin-left:24px}
@media(max-width:900px){.publication-heading{display:block}.publication-heading>.text-link{margin-top:25px}.publication{grid-template-columns:190px minmax(0,1fr);gap:30px}}
@media(max-width:600px){.publications{padding-top:55px}.publication{grid-template-columns:1fr;gap:18px;padding:28px 0}.publication h3{font-size:20px}.publication-meta{gap:5px}.publications-jump{margin-left:0;display:table}.publication-heading>.text-link{font-size:14px}.publication-heading{margin-bottom:28px}}

/* Real PDB coordinates rendered in 3D, with accessible motion controls. */
.molecule-figure{margin:0;min-width:0;padding:12px 0 10px 26px}
.molecule-label{display:flex;justify-content:space-between;gap:12px;font-size:13px;color:var(--muted)}
.pdb-id{font-size:12px;white-space:nowrap;letter-spacing:.06em}
.molecule-stage{height:440px;position:relative;margin:12px 0}
#molecule-viewer{position:absolute;inset:0;overflow:hidden;border-radius:4px;touch-action:pan-y}
#molecule-status{position:absolute;top:42%;left:10%;right:10%;text-align:center;font-size:14px;color:var(--muted);pointer-events:none}
#molecule-status[hidden]{display:none}
.molecule-controls{display:flex;flex-wrap:wrap;gap:8px;padding-bottom:18px;border-bottom:1px solid var(--line)}
.molecule-controls button{font:inherit;font-size:13px;min-height:38px;padding:7px 11px;border:1px solid #b8c4bc;border-radius:4px;color:var(--ink);background:transparent;cursor:pointer}
.molecule-controls button:hover{background:#e9ecdf}.molecule-controls button:disabled{opacity:.5;cursor:wait}
.molecule-controls button:focus-visible{outline:3px solid #7c9420;outline-offset:3px}
.molecule-legend{display:flex;gap:22px;flex-wrap:wrap;font-size:13px;margin-top:17px}
.molecule-legend span{display:flex;gap:8px;align-items:center}.molecule-legend i{width:10px;height:10px;border-radius:50%}.antibody-key{background:#12656a}.antigen-key{background:#a5be32}
#molecule-caption p{font-size:12px;color:var(--muted);margin:10px 0 4px}#molecule-caption>a{font-size:12px;color:var(--muted)}
@media(max-width:900px){.molecule-figure{padding-left:15px}.molecule-stage{height:360px}.molecule-label{flex-wrap:wrap}.molecule-controls{gap:6px}.molecule-controls button{font-size:12px;padding:7px 8px}}
@media(max-width:680px){.molecule-figure{padding:20px 0 0;max-width:none;width:100%}.molecule-stage{height:390px}.molecule-controls button{min-height:44px;font-size:13px}.molecule-label{font-size:14px}}

/* Bordered service cards, two across on desktop and stacked on mobile. */
.services{gap:24px;align-items:stretch}
.service{display:flex;flex-direction:column;min-width:0;padding:32px;border:1px solid #cfd8cf;border-radius:14px;background:#fffefa;box-shadow:0 5px 20px rgba(7,62,66,.035)}
.service-number{align-self:flex-start;display:inline-block;background:#edf2d7;border-radius:5px;padding:6px 10px;margin:0 0 26px;color:#526514;line-height:1.5}
.service>div{display:flex;flex-direction:column;flex:1;min-width:0}
.service h3{font-size:clamp(24px,2.2vw,30px);line-height:1.2}
.service details{margin:14px 0 22px}
.service .deliverable{margin-top:auto;padding-top:20px}
@media(max-width:760px){.services{gap:20px}.service{padding:26px;border-radius:12px}.service-number{margin-bottom:22px}.service h3{font-size:27px}}
@media(max-width:400px){.service{padding:23px}.service h3{font-size:25px}}

/* Founder portrait beside the biography. */
.about{align-items:center;gap:7%;grid-template-columns:minmax(0,.9fr) minmax(0,1.15fr)}
.about-portrait{margin:0;overflow:hidden;border-radius:14px;background:#e4e5db}
.about-portrait img{display:block;width:100%;height:auto;aspect-ratio:4/5;object-fit:cover}
.about-title .eyebrow{margin-bottom:20px}
.about-title .credentials{margin-bottom:30px;font-size:17px}
.about-title .credentials>span[aria-hidden]{margin:0 8px;color:#87958a}
@media(max-width:760px){.about{grid-template-columns:1fr;gap:32px}.about-portrait{width:100%;max-width:420px}.about-title .credentials{margin-bottom:24px}}